What would cause this weird bearing wear?

Daves69 said

I'm sure more experience will come. I'll ad this though.
I wouldn't be surprised on the mismatch from factory. But then, unless you've had the engine since manufactured, you likely don't know if the caps are original to the block I'd think.
It doesn't sound as though you are seeing or feeling rotation resistance due to the condition.
When you say "like one side I can catch my finger nail while I can't on the other" do you mean front to back or cylinder bank to cylinder bank?
Do the caps or block have corner breaks that can enhance the issue?
Did your main bolts turn in straight or wobble as you ran them down?
You could try to reset the caps. It may just end up the same, split the difference, or even move the offset to the opposite side.
The thrust bearing will control the cranks fore and aft motion.

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I think the mains are original, but even if they aren't they were line hones so they will work, and everything was measured so I'm sure they will work.
What I mean is that from cylinder bank to cylinder bank is slightly different on one main only, the rest are pretty close that way. The others slightly hang over going front to back, It was probably like this last time I put the motor together as well without ill effects, but my other bearing issues earlier in this thread were caused by the crank journal not being as round as it could be.

The bolts went in straight as well and, just dont want to have something slightly out of whack causing the bearing to be off, but then again that could just be the way the cap was originally when it was line honed? I will put up some pictures as it just makes things easier.