Drivetrain vibration

One thing that comes to mind is the torque converter.
If the engine was internally balanced, the torque converter should be too.
You may have gotten a converter that was meant for an externally balanced engine, but sold to you as an internally balanced one. (misboxed etc.)
The externally balanced one should have some big weights that look like 1/4'' x 1'' flat bar about an inch or so long welded to the side of it, (pretty easy to spot with the inspection cover off).
A neutrally balanced one should have no weights on it or very small ones.
It's pretty easy to knock them off with a good hammer and chisel.
Do you have a stock flex plate on it?
An aftermarket one might have weights on it or either of them could be bent.
I would contact your converter manufacturer to verify this information.............
Like your thoughts here, especially since the converter was in the car back when I first built it. I believe there is a weight on the outside of the housing. I will have to check more closely. I called TCI and they did mention the external/internal balancing, so you mentioning this makes me even more curious. I think the flex plate is original, and when bolting it up, each ear seemed even with the tabs on the converter. I will also check my pinion angles again and see if there is any runout on the driveshaft. I have a lift, so I can jack up the rear tires and run it through some gears.