1969 Race Dart w/ 426 Hemi- garaged for 45+ years

in the nuanced idea that the intake and motor go together and were probably purchased and raced that way until "upgraded" to the tunnel ram, sure.

the idea of several year old race motor in a year or two old car absolutely tracks with how things would play out back then. but another possibility, especially after seeing the motor broken down, is that it was purchased as parts, and put together with what all was available-- hence the alum rods, backyard balance job on the stock crank and the big ol gnarly cam. which again, under investigation of the trans, cut up floor, seats, and ladder bars kind of sort of tracks.

and as time rolled on, maybe your FIL collected the bits to make it more "SS" and less home brew?
The crossram was a super stock piece in 68 by the early 70’s to mid 70’s tunnel rams were used in pro stock on the darts for a couple years that included updates to the body to keep it legal, because the body was a 69 and not changed and the owner has a trophy from 72 the car was most likely run in modified production. Bracket racing was starting to happen but mostly populated at that time by cars that couldn’t run the numbers in class racing.