Ok this is a couple months old, but here goes:
I have a build that is very similar to yours. My goal is high performance daily driver.
I have a 1991 roller 360 with 10.2 compression with Lunati voodoo roller cam, a tuned Street Demon 750.
I have an A999 as well, but with the Lock-up totally removed. I was able to use Turbo action Cheetah stage 2 valve body and Hughes non-LU TQ converter.
(use at least 2500 stall!) I made the mistake and used a 2000 stall.
my rear gears is 3.21.
Yes there is a way to "shut-off " the lock-up or better yet do a mod similar that
@TrailBeast utilized. Especially with those 3.55 gears.
Couple of points:
3.55 gears with the A999 is still a very good all-around "fun" gear around town but will take some of the highway ability.
I'm sure your aware of the A999's 1 & 2 gear ratio vs. 727 and regular 904's. So with your combo your first gear take-off is equivalent to having a 727 with 3.96 rear gears.
With my 3.21 rear my first gear take-off is slightly higher vs. 727 with 3.55 rear. But I can cruise at 72 mph at 3000 rpm.
With your gears: (and I say this because I've had personal experience with using the same gears as you before)
With 26 inch rear tires at 70 mph my rpm's jumped to almost 3400 (something like 3370-3380 to be exact)
With 27 inch tires and 3.55 that would be a better match.
(I'm still thinking of going back to 3.55 gears Since I'm now going to 17 in rims with 255/50R17 rear tires which are 27 inches tall)
With the lock-up you're going to drop about 200+ rpms at highway cruise.
There are pros and cons to our setups:
My setup (LU totally removed/converted non-LU)
+ Cheaper Torque Converters
+ Likely stronger.
+ Less worry about electronics gadgets and such.
Keeping LU:
+ The trans runs cooler, especially at freeway speeds.
+ Flexibility in using a little higher (numerically) gear numbers.
+ You will gain slightly more mileage.
For example:
On my drive to Louiseville CO (from Ogden UT area) 500+ mile one way.
I usually get a best of around 17.5 mpg, but ranges can also average from 15.4 to 16.8 depending on crosswinds (Wyoming for you)
* That's maintaining 70-72 mph keeping it at 3000 rpm and under
* I did get almost about 18.2 mpg one time doing mainly 60-65 mph in the backroads (around 2500-2700 rpm) Sunny, clear 85 degree day with a modest 25mph tailwind.
* My best educated guess is you can gain anywhere between 3-5% increase in mileage (not factoring wind, temp, incline etc.) by keeping that lock-up function - Granted my math is nowhere as elaborate as
@AJ/FormS
* With an OD trannys like A500/42RH/46RH, that's a total game changer, expect about a 25-30% increase in mileage on hwy. due to their .69 OD gear.