You didn't specify a Leakdown result, so I might assume you did it at BDC, and that cannot work unless you back off all the rocker gear.
But you give a confident big fat zero for compression, and that also is not possible, unless maybe the intake is completely isolated from the atmosphere, but the comment "fires but won't start, precludes that. At least you didn't say backfires and lit the car on fire,lol. So in that case, at least the intakes are closing.
So far we know the pistons are going up and down, the intakes are closing, and somehow with zero compression, she fires but won't start.
Hmmmmmmmmmmmmmmmmmmmmmmmmmm
My guess is your gauges are lying,charge the battery, pull the plugs, squirt a lil oil in each hole, crank it over to distribute the oil and blow out the excess. Yeah it will make a mess so take steps to minimize that. Then repeat the compression test with a different screw-in tester, or do the LD test with the pistons at TDC-compression,each one in it's proper sequence. Or if you cannot get the pistons to hold still, sometimes it's just easier to do the LD test with the pistons at the bottom and the rocker gear backed off. Unfortunately, this does not tell us anything about the valve seal AFTER the rocker gear is re-installed; it only tells us if the chamber can or cannot be sealed, not is sealed.
Then post up the results.
BTW; some LD into the cc would be normal, say up to 2 or 3%, I mean the air will find the ring-gaps pretty quick